Peer Reviewed Articles

All downloads are in .pdf () format.

Dudzinski, K.M., Neff, N.A. 1990. A technique for the combination of clearing, staining, and injecting small mammals. Stain Technology 65(3):113-118.

Dudzinski, K.M., Frohoff, T.G., Shoda, L.K.M., Sparks, T.D. 1991. The Pine Warbler song repertoire: a preliminary description and analysis. Bulletin of the Texas Ornithological Society 24(2): 30-38.

Dudzinski, K.M., Frohoff, T.G., Crane, N. 1995. Behavior of a lone female bottlenose dolphin (Tursiops truncatus) with humans off the coast of Belize. Aquatic Mammals 21(2): 149-153.

Dudzinski, K.M., Clark, C.W., Würsig, B. 1995. A mobile video/acoustic system for simultaneously recording dolphin behavior and vocalizations underwater. Aquatic Mammals 21(3): 187-193.

Dudzinski, K.M. 1998. Contact behavior and signal exchange among Atlantic spotted dolphins (Stenella frontalis). Aquatic Mammals 24(3): 129-142.

Dudzinski, K.M., M. Sakai, M., Masaki, K., Kogi, K., Hishii, T., Kurimoto, M. (2003). Behavioral observations of adult and sub-adult dolphins towards two dead bottlenose dolphins (one female and one male). Aquatic Mammals 29(1): 108-116.

Kogi, K., Hishii, T., Imamura, A., Iwatani, T., Dudzinski, K.M. (2004) Preliminary demographic parameters and reproductive rates of Indo-Pacific bottlenose dolphins (Tursiops aduncus) around Mikura Island, Japan. Marine Mammal Science 20(3): 510-526.

Paulos, R.D., Dudzinski, K.M., Kuczaj, S.A. 2007. The role of touch in select social interactions of Atlantic spotted dolphin ( Stenella frontalis ) and Indo-Pacific bottlenose dolphin ( Tursiops aduncus ) Ethology (Online First) (340k)

Gregg, J.D., Dudzinski, K.M., Smith, H.V. (2007) Do dolphins eavesdrop on the echolocation signals of conspecifics? International Journal of Comparative Psychology, 20: 65-88 (170kb)

Gregg, J.D., Dudzinski, K.M., Smith, H.V. (2007) 3D MASC: a method for estimating relative head angle and spatial distance of dolphins from underwater video footage. Animal Behaviour doi:10.1016/j.anbehav.2007.09.009 (352k)

Dudzinski, K.M., Thomas, J. Gregg, J.D.(2008) Communication. In (W.F. Perrin, B. Würsig, H.C.M. Thewissen, eds) Encyclopedia of Marine Mammals, second edition. Academic Press, Inc.

Dudzinski, K.M., Gregg, J.D., Ribic, C.A., Kuczaj, S.A. (2009) A comparison of pectoral fin contact between two different wild dolphin populations. Behavioural Processes, 80:182-190

Dudzinski, K.M., Gregg, J.D., Paulos, R.D., Kuczaj, S.A. (2010) A comparison of pectoral fin contact behaviour for three distinct dolphin populations. Behavioural Processes, 84:559–567 (Contact DCP (info{at}dcpmail{dot}org) to request a PDF)

Dudzinski, K.M., S.J. Brown, M. Lammers, K. Lucke, D. Mann, P. Simard, C. Wall, M.H. Rasmussen, E.E. Magnúsdóttir, J. Tougaard, and N. Eriksen. 2010. Trouble-shooting deployment and recovery options for various stationary passive acoustic monitoring devices in both shallow and deep water applications. Journal of the Acoustical Society of America Vol. 129, Issue 1 (2MB)   (Abstract: Copyright (year) Acoustical Society of America. This article may be downloaded for personal use only. Any other use requires prior permission of the author and the Acoustical Society of America.)

Greene, W., Melillo-Sweeting, K., Dudzinski, K. International Journal of Comparative Psychology 24(3):292-306. 2011. Comparing object play in captive and wild dolphins.

Melillo-Sweeting, K., Reid, J., Gittens, L., Adimey, N., Dillet, J. Observations and relocation of a West Indian manatee (Trichechus manatus) off Bimini, The Bahamas. Aquatic Mammals 37(4): 502 - 505. DOI 10.1578/AM.37.4.20011.502. Visit to purchase or Contact DCP (info{at}dcpmail{dot}org) to request a PDF.

Dudzinski, K.M., Gregg, J.D., Melillo-Sweeting, K., Seay, B., Levengood, A., Kuczaj, S.A. (2012) Tactile Contact Exchanges Between Dolphins:  Self-rubbing versus Inter-individual Contact in Three Species from Three Geographies. International Journal of Comparative Psychology, 2012, 25, 21-43

Vaughn-Hirshorn, R., Hodge, K.B., Wursig, B., Sappenfield, R.H., Lammers, M.O., Dudzinski, K.M. (2012). Characterizing dusky dolphin sounds from Argentina and New Zealand. Journal of the Acoustical Society of America. 132(1): 498-506. Visit or Contact DCP (info{at}dcpmail{dot}org) to request a PDF.

K. M. Dudzinski, C. D. Marshall, & J. S. Reidenberg. 2013. Air Release from the Left Orbit of an Indo-Pacific Bottlenose Dolphin (Tursiops aduncus): Symptomatic and Anatomical Aspects. Aquatic Mammals 39(1): 97-100.

Whitt AD, Dudzinski K, and Laliberté JR. 2013. North Atlantic right whale distribution and seasonal occurrence in nearshore waters off New Jersey, USA, and implications for management. Endangered Species Research 20:59-69. To request a PDF copy of this article, please email Amy Whitt at This email address is being protected from spambots. You need JavaScript enabled to view it.

Vaughn-Hirshorn, R.L., Muzi, E. Richardson, J.L., Fox, G.J., Hansen, L.N., Salley, A.M., Dudzinski, K.M., Würsig, B. (2013) Dolphin underwater bait-balling behaviors in relation to group and prey ball sizes, Behavioural Processes, I7, 10.1016/j.beproc.2013.04.003.

Evans-Wilent, J, Dudzinski, KM. (2013) Vocalizations associated with pectoral fin contact in bottlenose dolphins (Tursiops truncatus). Behavioural Processes, 100, 74– 81. Contact DCP for a PDF.

Melillo-Sweeting, K, Turnbull S.D. and Guttridge, T.L. (2013). Evidence of shark attacks on Atlantic spotted (Stenella frontalis) off Bimini, The Bahamas. Marine Mammal Science 30(3):1158-1164. DOI:10.1111/mms.12082. 

Melillo-Sweeting, K., Yeater, D., Dudzinski, KM. (2015). Dolphin sightings near the coast of Bimini, The Bahamas, 2003 – 2013. Aquatic Mammals 41.3: 245-251. DOI: 10.1578/AM.41.3.2015.245. Visit to purchase or contact DCP (info{at}dcpmail{dot}org) to request a PDF.

Harvey, B.S., Dudzinski, K.M., Kuczaj, S. A. (2017). Associations and the role of affiliative, agonistic, and socio-sexual behavior among common bottlenose dolphins (Tursiops truncatus). Behavioural Processes. 135: 145-156. 

Dudzinski, KM, Ribic, CA (2017). Pectoral fin contact as a mechanism for social bonding among dolphinsAnimal Behavior & Cognition 4(1): 31-49.

Kaplan, JD, Melillo-Sweeting, K, Reiss, D. (2017). Biphonal calls in Atlantic spotted dolphins (Stenella frontalis): bitonal and burst-pulse whistles. Bioacoustics. DOI:10.1080/09524622.2017.1300105. 

Davis GE, Baumgartner MF, Gurnee J, Bell J, Berchok C, Bort Thornton J, Brault S, Buchanan G, Charif RA, Cholewiak D, Clark CW, Corkeron P, Delarue, Dudzinski K, Hatch L, Hildebrand J, Hodge L, Klinck H, Kraus S, Martin B, Mellinger D, Moors-Murphy H, Nieukirk S, Nowacek D, Parks S, Read A, Rice AN, Risch D, Širović A, Soldevilla M, Stafford K, Stanistreet J, Summers E, Todd S, Warde A, Van Parijs SM (2017) Long-term passive acoustic recordings track the changing distribution of North Atlantic right whales (Eubalaena glacialis) from 2004 to 2014. Scientific Reports, 7: 13460. DOI:10.1038/s41598-017-13359-3. 

Eierman, LE, Laccetti, K, Melillo-Sweeting, K, Kaplan, JD. (2019). Interspecies pectoral fin contact between bottlenose dolphins and Atlantic spotted dolphins off Bimini, The Bahamas. Animal Behaviour 157: 167-176. Visit Animal Behaviour or Contact DCP (info{at}dcpmail{dot}org) to request a PDF.

Kaplan, JD. Goodrich, SY, Melillo-Sweeting K, Reiss, D. (2019). Behavioural laterality in foraging bottlenose dolphins (Tursiops truncatus). Royal Society Open Science. 6: 190929.


Book Chapters

Dudzinski, K.M., Douaze, E., Thomas, J. (2002) Communication. In (W.F. Perrin, B. Würsig, H.C.M. Thewissen, eds) Encyclopedia of Marine Mammals. Academic Press, Inc.

Dudzinski, KM, Gregg, J (2017). Communication. In (W.F. Perrin, B. Würsig, H.C.M. Thewissen, eds) Encyclopedia of Marine Mammals, third edition. Academic Press, Inc. (pp. 210-215).

Dudzinski, KM, Hill, HM (2018). Cetacean Communication. Pp. 1-12. In (J Vonk, TK Shackelford eds). Encyclopedia of Animal Cognition and Behavior. Springer International Publishing AG. DOI: 10.1007/978-3-319-47829-6_991-1

Popular Publications (sample only)

Dudzinski, K.M., Graham, C.M. (1992). An Introductory Trail Guide to Blackbird Caye. Texas A&M University Print Shop, Galveston. 62pp.

Dudzinski, K.M., Fertl, D., Megill, W., Baird, R.W., Stern, S.J. (1995). Scientific correspondence: Report on group size workshop from the Tenth Biennial Conference on the Biology of Marine Mammals. Email: Marmam Net. 3pp.

Dudzinski, K.M. (1996). Final Summary Report to OSE: Communication in Atlantic Spotted Dolphins (Stenella frontalis): Relationships between vocal and behavioral activities. Oceanic Society Expeditions. San Francisco. 18pp.

Dudzinski, K.M., Perrine, D. (1997). Dolphins Behaving Badly? Dive International Magazine. December Issue. Amano, M. Barbosa, C., Dudzinski, K.M., Kawaguchi, N., Kikuchi, K., Mori, Moyer, J.T., K., Orams, M.C., Soeda, R., Taguchi, S., Yoshioka, M. 1998. Interactions with Dolphins and Whales. Japanese Publishers & sponsored by Omega.

Dudzinski, K. M. (1998). The Best Kept Secret in Dolphin Swim Programs. Whalewatcher. Fall/Winter. 6 pp.

Dudzinski, K.M. (2003) Minding Our Manners: Learning the proper etiquette for Swimming with Dolphins by Observing their Interactions. In: TG Frohoff, B Petersen (editors). Between Species. Sierra Club Books.

Gregg, J. (2012). Will humans ever be able to speak or even understand dolphin? New Scientist, July 7, 57.

Melillo-Sweeting, K. (2018). Connect the Dots. Coastal Angler Magazine, January.


Dudzinski, K.M. (2000) Meeting Dolphins: My adventures in the Sea. National Geographic Publications.

Dudzinski, K.M., Frohoff, T. (2008) Dolphin Mysteries: Unlocking the secrets of communication. Yale University Press.

Gregg, J. (2013) Are dolphins really smart? The mammal behind the myth. Oxford: Oxford University Press.

Theses and Dissertations

Reily, G. (2014) Whistle Types in a Captive Bottlenose Dolphin Population (Tursiops truncatus). Unpublished Master's Thesis. NOVA Southeastern University, Florida.

Levengood, A. (2013) Is blood thicker than water? The role of kin and non-kin in non-mother-calf associations of captive bottlenose dolphins (Tursiops truncatus).Unpublished Master's Thesis. St. Andrews University, Scotland.

van Hoey, Ni. (2013) Photo - identification and distribution of bottlenose dolphins ( Tursiops truncatus) off Bimini, The Bahamas, 2006 - 2009. Unpublished Master's thesis, Alaska Pacific University, Anchorage, USA

McIntosh, B. (2013) Examining dolphin whistles via neural networks. Unpublished Master's Thesis. SUNY Buffalo, NY.

Miller, L. (2010) Conflict management in Indo-Pacific bottlenose dolphins. Unpublished Master's Thesis. University of Southern Mississippi, Hattiesburg, MS.

Beard, K. (2008) Use of Bubble Emissions by Bottlenose (Tursiops truncatus) and Atlantic Spotted Dolphins (Stenella frontalis) Relative to Age and Sex. Unpublished Master's Thesis. The University of Connecticut, Storrs, CT.

Gregg, J.D. (2008) Joint Attention and Echoic Eavesdropping in Wild Bottlenose Dolphins (Tursiops aduncus). Unpublished doctoral dissertation, University of Dublin, Trinity College, Dublin, Ireland.

Makepeace, S. (2008) Structural changes in dolphin tonal vocalizations in a changing soundscape: A field study. Unpublished Master's Thesis

Melillo, K. (2008) Interspecific interactions between Atlantic spotted (Stenella frontalis) and bottlenose dolphins (Tursiops truncatus) off Bimini, Bahamas 2003 – 2007. Unpublished Master's thesis, Alaska Pacific University, Anchorage, USA. 68 pp

Paulos, R. (2004) Non-vocal communication in the Atlantic spotted dolphin (Stenella frontalis) and the bottlenose dolphin (Tursiops truncatus): sequential analyses of inter- and intra-specific patterns of behavior. Unpublished Master's Thesis, College of Education and Psychology of The University of Southern Mississippi

Dudinski KM (1996) Communication and behavior in the Atlantic spotted dolphins (Stenella frontalis): relationship between vocal and behavioral activities. Dissertation Thesis, Texas A&M University, College Station.


Scientists Collaborating with DCP

Hendry, J. L. (2004). The ontogeny of echolocation in an atlantic bottlenose dolphin (Tursiops truncatus). Unpublished Doctoral Dissertation, College of Education and Psychology of The University of Southern Mississippi. (820kb)

Blomqvist, C., Amundin, M. High-Frequency Burst-Pulse Sounds in Agonistic/Aggressive Interactions in Bottlenose Dolphins, Tursiops truncatus. In: Echolocation in Bats and Dolphins. The University of Chicago Press. (1.1 MB)

Blomqvist, C., Amundin, M. 2004. An Acoustic Tag for Recording Directional Pulsed Ultrasounds Aimed at Free-Swimming Bottlenose Dolphins (Tursiops truncatus) by Conspecifics. Aquatic Mammals 30(3): 345-356. (1.7 MB)

Blomqvist, C., Mello, I., Amundin, M. 2005. An Acoustic Play-Fight Signal in Bottlenose Dolphins (Tursiops truncatus) in Human Care. Aquatic Mammals 31(2): 187-194 (3.9 MB)

Kuczaj, S.A. & Yeater, D.B. (2006). Dolphin imitation: Who, what, when, and why? Aquatic Mammals, 32, 413-422 . (116 kb)

Highfill, Lauren E.; Kuczaj II, Stan A. Do Bottlenose Dolphins (Tursiops truncatus) Have Distinct and Stable Personalities? Aquatic Mammals, Vol. 33, No. 3. (September 2007), pp. 380-389. (117 kb)

Kuczaj, S.A. II & Walker, R.T. (2006). How do dolphins solve problems? In E.A. Wasserman & T.R. Zentall (Eds.), Comparative Cognition: Experimental Explorations of Animal Intelligence. New York: Oxford University Press. pp. 580-600. (2.27 MB)

Independent Study Paper written in conjunction with DCP

Lovejoy, P. (2013) Examining Social Quality between Dolphin Dyads. Independent Study Paper. Connecticut College: New London, CT

Contact Us

Write to us via snail-mail at:

Dolphin Communication Project
P.O. Box 7485
Port St. Lucie, FL, 34985

Email us:

info {at} dcpmail {dot} org


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